Date of the first record

The date of the first documented record of the species occurrence in a country/country region.
Date fields usage example:

Date to be specifiedDate FromDate To
Exact 198519851985
18th century17011800
before 1700 1700
after 20012001 

Environmental position

Environment(s) occupied by a species throughout its life cycle.

OPTIONS:

Biofouling. Assemblage of organisms on wetted artificial substrates.

Commensal. An organism in a symbiotic relationship, in which one benefits while the other is not adversely affected.

Demersal. Synonyms: hyperbenthic, benthopelagic, nektobenthic. An organism living at, in or near the bottom of the sea, but having the ability to swim.

Ectoparasite. A parasite living on the surface of its host.

Endoparasite. A parasite living within the organs or tissues of its host.

Epifaunal. Synonym: epibenthic. An animal inhabiting the surface of the seabed, submerged plants and animals.

Epilithic. An organism living on the surface of rock or other hard inorganic substrata.

Epiphytic. An organism living on the surface of a plant, non-parasitic.

Epizoic. An organism living on the surface of an animal, non-parasitic.

Infaunal. Synonym: endobenthic. An animal living within the seabed sediments.

Interstitial. An organism (< 1 mm) living in the spaces between sediment particles.

Lithotomous. An organism burrowing into rock.

Neustonic. An organism living on (epineuston) or under (hyponeuston) the surface film of water bodies.

Pelagic. An organism inhabiting the water column.

Pleustonic. An organism inhabiting the water surface due to their own buoyancy, normally positioned partly in the water and partly in the air.

Habitat type

Estuary. River mouth, transition zone between riverine and marine environments, subject to influences from both.

Lagoon. Shallow, enclosed water body separated from the sea by barrier islands, narrow spit or reefs.

Offshore. Areas located at least 50 nautical miles from the shore.

Open coast. A coast not sheltered from the sea.

Strait/Sound. Channels between the mainland and an island or between two islands which are open at both ends to the open coast (it does not refer to similar features or narrows within marine inlets).

Sheltered coastal area. Coastal area partly surrounded by land (e.g., bay, inlet, fjord).

Ports. A location on a coast or shore containing one or more harbours where ships can dock and transfer people or cargo to or from land.

Port vicinity. The area near a port where ballast water operations may occur, including areas where vessels may conduct ballast water discharge or uptake operations when approaching a port or leaving it, e.g., port approaches, anchorage areas and designated ballast water exchange areas. The dimension is port specific.

Marina. A specially designed harbour for pleasure craft and small boats.

Aquaculture sites. Areas set out for the purpose of farming aquatic organisms.

Marine Protected Area (MPA). Defined marine area where natural resources are given greater protection than the surrounding waters. Different categories exist depending on the level of protection afforded by legislation.

Migration pattern

Diurnal. Movements between alternative habitats over day and night, e.g. vertical migration.

Life-time. One time migration between different habitats during the life cycle, e.g. anadromus migration.

Not relevant. No evidence of any life history cycle stages to migrate.

Seasonal. Movements between alternative habitats during a specific time of a year (e.g., spawning and feeding migrations).

Pathway / Vector

Pathway

A pathway is the route a NIS takes to enter or spread through a non-native ecosystem e.g. vessels. Each pathway may have a number of vectors.

Vector

A vector is a transfer mechanism and is the physical means by which species are transported from one geographic region to another. More than one vector within a pathway may be involved in a transfer of species.

Pathways and vectors included:
PathwayVector
Aquarium tradeIntentional organism release
Transported water
Waste discharge
Culture activitiesAquaculture equipment
Associated water & packaging material
Intercontinental stock movement
Regional stock movement
Unintentional release & escapees
Leisure activitiesAngling catch
Cultural releases
Live bait
Live souvenirs
Sport equipment
Stocking for angling
Waste discharge
Live food tradeIntentional organism release
Transported water
Waste discharge
ManagementBiological habitat management
Construction equipment
Construction materials
Release for biological control
Natural spread from neighboring countriesOther natural vectors
Water currents
Other canalsCanal de Midi (linking the Bay of Biscay with the Mediterranean Sea)
Kiel Canal (linking the North Sea with the Baltic)
Northern waterway (linking the Baltic with the Ponto-Caspian region through Volga river canal system)
Rhone waterway (linking the North Sea with the Mediterranean)
Southern waterway (linking the North Sea with the Black Sea through Danube river canal system)
Central waterway (linking the Baltic Sea with the Black Sea through the Dnieper river canal system)
Other waterways
Irrigation canals
Research and educationGear movement
Intentional releases
Unintentional release & escapees
Waste discharge
Suez Canal 
VesselsAnchor and anchor chain
Ballast tank sediments
Ballast water
Ship’s hull
Sea chest
Others
Wild fisheriesDiscard of by-catch
Fishing gear
Live bait release
Live packaging material
Processed live material
Stock movements
Transported water

Pathway / Vector – Levels of certainty:
LevelCriteriaExamples
Direct evidenceThe species was actually found associated with the specific vector(s) of a pathway at the time of introduction to a particular locality within a country/country region.Documented evidence of an introduction: release to the wild for stocking or biological control; escape/release of live food; import of cultured species and documented findings of their associate organisms, parasites and diseases on transmission; appearance of organisms by hull fouling, ballast water discharge sampling or other ship vectors documented upon an arrival with appropriate scientific methods.
Very likelyThe species appears for the first time in a locality where a single pathway/vector(s) is known to operate and where there is no other explanation that can be argued for its presence except by this likely pathway/vector(s).A highly localized distribution of a species in an area adjacent to an isolated port or other locality where the only pathway is vessels and its vector(s) (ballast water, hull fouling, etc). This often involves geographically discontinuous distributions. It may be a continuous spread as in case of introduction by canals or by natural means. The conclusion is deduced from the analysis of the invasion event and species distribution patterns.
PossibleThe species cannot be convincingly ascribed to a single pathway, but is known to be introduced by this pathway(s) elsewhere.There may be more than one pathway involved in the introduction within a country/country region. Arrival of a species known to have taken place elsewhere by the same pathway(s) which operates in an area. A conclusion is made by expert judgment based on pathways currently or historically present.
UnknownInvasion of a given alien species cannot be clearly explained.Where no rational explanation for the appearance of a species in a given country/region.

Population status

Population status (the lowest level of certainty):

Unknown. There is no reliable information on population status of a species.
Established. A species is known to form a reproducing population in a wild.
Not established. There is no evidence of a species’ reproducing population in a wild.

Population status (the moderate level of certainty):
Extinct/no recent record. There are old records where a species was recorded but have not been seen in the same region since.
Rare/single record. There are only casual observations or a single record of a species'presence available.
Common. A species with successfully reproducing populations in an open ecosystem, which are unlikely to be eliminated by man or natural causes. Not dominating native communities.
Abundant. A species with successfully reproducing populations in an open ecosystem, which are unlikely to be eliminated by man or natural causes. Locally dominating native communities.
Very abundant. A species with successfully reproducing populations in an open ecosystem, which are unlikely to be eliminated by man or natural causes. Largely dominating native communities.
Outbreak. A species undergoing pulse-like, short-term (days to few months) exponential population growth during which they have an adverse effect on one or more of the following: biological diversity, ecosystem functioning, socio-economic values and human health.

Recipient region

The country/region for which introduction is recorded.

References.



References should follow the standard of Biological invasions:


Journal article
Gamelin FX, Baquet G, Berthoin S, Thevenet D, Nourry C, Nottin S, Bosquet L (2009) Effect of high intensity intermittent training on heart rate variability in prepubescent children. Eur J Appl Physiol 105:731-738. doi: 10.1007/s00421-008-0955-8
Ideally, the names of all authors should be provided, but the usage of “et al” in long author lists will also be accepted:
Smith J, Jones M Jr, Houghton L et al (1999) Future of health insurance. N Engl J Med 965:325–329


Article by DOI


Slifka MK, Whitton JL (2000) Clinical implications of dysregulated cytokine production. J Mol Med. doi:10.1007/s001090000086


Book
South J, Blass B (2001) The future of modern genomics. Blackwell, London


Book chapter
Brown B, Aaron M (2001) The politics of nature. In: Smith J (ed) The rise of modern genomics, 3rd edn. Wiley, New York, pp 230-257


Online document
Cartwright J (2007) Big stars have weather too. IOP Publishing PhysicsWeb. http://physicsweb.org/articles/news/11/6/16/1. Accessed 26 June 2007


Dissertation
Trent JW (1975) Experimental acute renal failure. Dissertation, University of California

Reproductive duration

Long. Breeds in one or more discrete periods, each longer than three months.

Medium. Breeds in one or more discrete periods, each longer than a week and less than three months.

Short. Breeds in one or more discrete periods within a week.

Reproductive seasonality

Months for a species known to reproduce in the invaded site.

Salinity range

The exact salinity range if known (psu), else salinity zone(s) according to the Venice system:
1. Limnetic [<0.5psu]
2. β-Oligohaline [0.5-3psu]
3. α-Oligohaline [3-5psu]
4. β-Mesohaline [5-10psu]
5. α-Mesohaline [10-18psu]
6. Polymixohaline [18-30psu]
7. Euhaline [30-40psu]
8. Hypersaline [>40psu]

Source region

The area the species was introduced from to the recipient country/country region. Depending on the information availability may be ascribed to a particular locality (e.g. port), a country, a LME or a larger Ocean region.
CAUTION: in many cases the source area will be not the same as the area of native origin which is defined in the SPECIES block of the database.

Species status

Non-indigenous species. Non-indigenous species (synonyms: alien, exotic, non-native, allochthonous, introduced) are species, subspecies or lower taxa (such as a variety, form) introduced outside of their natural range (past or present) and outside of their natural dispersal potential. This includes any propagule of a NIS, such as a gamete, seed or resting spore, a gravid female or a pair of individuals of different sexes (in sexual reproduction), or a vegetative reproductive organ and section of tissue (in asexual reproduction), which might survive, reproduce and subsequently form a population. It also includes hybrids between an alien species and an indigenous species, fertile polyploid organisms and artificially hybridized species irrespective of their natural range or dispersal potential.

Cryptogenic. Cryptogenic species are such species which cannot be reliably demonstrated as being either introduced or native. In some cases the true origin of a species remains obscure because of either insufficient taxonomic knowledge or due to a lack of records from the time they became introduced, or for other reasons.

Temperature range

Indicate min. and max. annual temperature range in the area where a species is known to maintain an established (reproducing) population.

Wave exposure

Exposed. Open coastline facing prevailing wind and receiving both wind-driven waves and swell.

Semi exposed. Generally open coasts facing away from prevailing winds or sheltered by offshore reefs/structures.

Sheltered. Coasts with a restricted fetch (<20 km) and lacking persistent swell.

Zonation

Ecological zone(s) occupied by a species throughout its life cycle.

Benthic - Bathyal. Synonym: continental slope. The seafloor between the edge of the continental shelf and abyssal plain (200-4000 m).

Benthic - Littoral. Synonym: intertidal. The shore between the high and low water marks.

Benthic - Sublittoral beyond photic zone. Synonym: lower circalittoral. The lower part of the continental shelf, where photosynthesis cannot take place.

Benthic - Sublittoral within photic zone. Synonyms: subtidal, infralittoral. The shallow part of sublittoral where photosynthesis can occur.

Benthic - Supralittoral. Synonyms: splash zone, spray zone, supratidal zone. The area above the spring high tide line, subject spray or splash.

Pelagic - Littoral. Water mass within littoral zone.

Pelagic - Neritic. Water mass above the continental shelf.

Pelagic - Offshore. Synonym: oceanic. Water mass beyond the continental shelf.

Public domain: Introduction event account

Species Austrominius modestus  
Date of the first record (?) 1957

References (not structured):
Beard DM (1957) Occurrence of Elminius modestus Darwin in Ireland. Nature 180: 1145.

Comments:
It was first located in lough hyne on the south coast of Ireland during September 1957. This region was a special marine study area in Ireland, although not a reserve at this time.
Recipient region (?) Country: Ireland
LME: 24. Celtic-Biscay Shelf
LME sub-region: Celtic seas


References (not structured):
Beard DM (1957) Occurrence of Elminius modestus Darwin in Ireland. Nature 180: 1145.

Comments:
In the region of Lough Hyne specimens were found close by in Tralispean and on the Coosh at the entrance to the Lough.
Source region (?) Not entered

References (not structured):
Crisp, D.J. 1958. The spread of Elminius modestus Darwin in north-east Europe. Journal of the Marine Biological Association of the United Kingdom, 37: 438-520
Pathway / Vector (?) Level of certainty: Highly likely

Pathway: Vessels
Vector: (Highly likely) Ship’s hull

References (not structured):
Bishop, M.W.H. 1947. Establishment of an immigrant barnacle in British coastal waters. Nature, 159: 501.

Comments:
The most generally accepted pathway is that of vessels and the vector the hull. Barnacles are frequently found associated with the hulls of vessels and even small craft such as dinghies. Its appearance of the south coast of Ireland has been associated with ports and harbours.
Habitat type (?) Aquaculture sites
Estuary
Lagoon
Marina
Port vicinity
Ports
Sheltered coastal area

References (not structured):
Allen, B.M., Power, A.M., O'Riordan, R.M., Myers, A.A. & McGrath, D. ; (2006) Increases in the abundance of the invasive barnacle Elminius modestus Darwin in Ireland. Biology and Environment, 106 (B):91-97.
Minchin D, Nunn J (2013) Rapid assessment of marinas for invasive alien species in Northern Ireland. A report undertaken for the Northern Ireland Environment Agency, Belfast. 96pp.

Comments:
Has been known to cause extensive fouling on aquaculture equipment and has caused rafts holding oysters to sink. Commonly associated with hull fouling especially on craft that are moored inter-tidally. Known from floating pontoons and piles at marinas.
Wave exposure (?) Sheltered

References (not structured):
Allen, B.M., Power, A.M., O'Riordan, R.M., Myers, A.A. & McGrath, D. (2006) Increases in the abundance of the invasive barnacle Elminius modestus Darwin in Ireland&. Biology and Environment, 106 (B):91-97.

Comments:
Generally the species occurs in sheltered areas most often in estuaries where it can be very abundant.
Salinity range (?) Venice system:
5. α-Mesohaline [10-18psu]
6. Polymixohaline [18-30psu]
7. Euhaline [30-40psu]

References (not structured):
O'Riordan RM, Murphy O (2000) Variation in the reproductive cycle of Elminius modestus in southern Ireland. Journal of the Marine Biological Association of the United Kingdom 80: 608-616.

Comments:
nauplius stages can endure salinities down to 20/21psu
Temperature range (?) Not entered
Zonation / Substratum (?) Benthic:
Littoral (Benthic)
Sublittoral within photic zone
Substratum:
Artificial (manmade)
Biogenic (living or nonliving)
Hard (cobbles to bedrock)
Soft (mud to pebbles)

References (not structured):
Allen, B.M., Power, A.M., O'Riordan, R.M., Myers, A.A. & McGrath, D. ; (2006) Increases in the abundance of the invasive barnacle Elminius modestus Darwin in Ireland. Biology and Environment, 106 (B):91-97.

Comments:
Attaches to a wide range of substrata including Littorina littorea in salt-marshes
Reproductive duration (?)Long

References (not structured):
Watson DI, O'Riordan RM, Barnes DKA, Cross T (2005) Temporal and spatial variability in the recruitment of barnacles and the local dominance of Elminius modestus Darwin in SW Ireland. Science (New York, N.Y.), 63 :119-131.
O'Riordan RM, Murphy O (2000) Variation in the reproductive cycle of Elminius modestus in southern Ireland. Journal of the Marine Biological Association of the United Kingdom 80: 608-616.

Comments:
The species is an obligatory cross-fertilising haemaphrodite which can produce multiple broods. well nourished adults produce broods at 8-31C. Larvae hatch at salinities above 21 psu and are then active between 19 and 40 psu.
Reproductive seasonality (?) Apr.
May
Jun.
Jul.
Aug.
Sept.
Oct.
Nov.
Dec.

References (not structured):
Watson, DI,O'Riordan, RM,Barnes, DKA,Cross, T; (2005) Temporal and spatial variability in the recruitment of barnacles and the local dominance of Elminius modestus Darwin in SW Ireland. Science (New York, N.Y.), 63 :119-131.
Harms, J. 1984. Influence of water temperature on larval development of Elminius modestus and Semibalanus balanoides (Crustacea, Cirripedia). Helgoländer Meeresuntersuchungen 38: 123-134.
O'Riordan RM, Murphy O (2000) Variation in the reproductive cycle of Elminius modestus in southern Ireland. Journal of the Marine Biological Association of the United Kingdom 80: 608-616.

Comments:
reproduction commences in the spring once temperatures exceed 6-8C
Migration pattern (?) Not entered
Population status (?) Very abundant (Moderate level of certainty)

References (not structured):
Allen, B.M., Power, A.M., O'Riordan, R.M., Myers, A.A. & McGrath, D. ; (2006) Increases in the abundance of the invasive barnacle Elminius modestus Darwin in Ireland. Biology and Environment, 106 (B):91-97.
Holmes JMC (1980) Some crustacean records from Lough Ine, Co Cork. Irish Biogeographical Bulletin 4: 33-37.
Watson DI, O'Riordan RM, Barnes DKA, Cross T (2005) Temporal and spatial variability in the recruitment of barnacles and local dominance of Elminius modestus Darwin in SW Ireland. Estuarine, Coastal and Shelf Science 63: 119-131.
Lawson J, Davenport J, Whitaker A (2004) Barnacle distribution in Lough Hyne Marine Nature Reserve: a new baseline and an account of invasion by the introduced Australasian species Elminius modestus Darwin. Estuarine, Coastal and Shelf Sciences 60: 729-735.
Crisp DJ, Southward AJ (1959) The further spread of Elminius modestus to the British Isles. Journal of the Marine Biological Association of the United Kingdom 38: 429-437.
King PA, Keogh E, McGrath D (1997) The current status of the exotic barnacle Elminius modestus Darwin in Galway Bay, Ireland. Irish Naturalists' Journal 25(10): 353-392.
McGrath D and King P (1992) The immigrant barnacle, Elminius modestus Darwin, in Galway Bay, Ireland.
Bulletin of the Irish Biogeographical Society 15: 63-67.
Aitken JJ, Hillis JP (1959) The occurrence of Elminius modestus (Darwin) in north Ireland. Irish Naturalists' Journal 13(4): 100.
O'Riordan RM (1996) The current status and distribution of
the Australian barnacle Elminius modestus Darwin in Ireland. In: Proceedings of the Irish Marine Science Symposium 1995 (eds) Keegan BF and R O'Connor, Galway University Press, Galway, 207-218.
O'Riordan, R.M.; (1996) 'The current status of the Australian barnacle Elminius modestus Darwin in Ireland' In: B.F. Keegan and R. O'Connor (eds). Proceedings of the Irish Marine Science Symposium 1995. Ireland: NUIG.
Minchin D, Nunn J (2013) Rapid assessment of marinas for invasive alien species in Northern Ireland. A report undertaken for the Northern Ireland Environment Agency, Belfast. 96pp.

Comments:
The species is locally abundant in estuaries such as Kinsale Estuary and is now present within most estuaries about most coastal areas. The barnacle was not found within the Lough HYne until it was recorded by Holmes (1980)at the North Pier. This Lough has since become a special site of study. While this was the first place where this barnacle became recognised it was most probably present in Cork Harbour between 1954 and 1956, it being absent when surveyed in 1953.(Crisp, 1958)
Species status (?) Non-indigenous species

References (not structured):
Pope EC (1966) Sessile barnacles Thoracica, Cirripedia: Port Phillip Survey 1957-1963. Memoirs of the National Museum. Victoria 27: 179-180.

Comments:
The species is native to Australia (and New Zealand). There is a further species that prefers more open coastal conditions, lower surfaces of boulders and in caves Elminius simplex
Port vicinitiesPort vicinity #1/14: Bantry
Date of the first record: 1968
Population status: Established
References:
Crapp GB (1973) The distribution of animals and plants on the shores of Bantry Bay. Scientific Investigations of the Fisheries Branch, Ireland, B, No 9, 1-35.
Baker JM, Hiscock S, Levell D, Bishop G, Precious M, Collinson R, Kingbury R, Sullivan AJ (1981) The rocky shore biology of Bantry Bay: a re-survey. Irish Fisheries Investigations, Series B. No 23: 1-27.
Comments:
The species is presently well established in Bantry Bay.

Port vicinity #2/14: Belfast
Date of the first record: 1958
Population status: Very abundant
References:
Crisp DJ, Southward AJ (1959) The further spread of Elminius modestus in the British Islea to 1959. Journal of the Marine Biological Association of the United Kingdom 64: 625-635.

Port vicinity #3/14: Castletownbere
Date of the first record: 1996
Population status: Not entered
References:
Minchin A, Minchin D (1997) Dispersal of TBT from a fishing port determined using the dogwhelk Nucella lapillus as an indicator. Environmental Technology, 18(12): 1225-1234
Comments:
Found during a survey attached to dogwhelk Nucella lapillus during July 1996.

Port vicinity #4/14: Cork
Date of the first record: 1958
Population status: Established
References:
Crisp DJ, Southward AJ (1959) The further spread of Elminius modestus in the British Islea to 1959. Journal of the Marine Biological Association of the United Kingdom 64: 625-635.
Comments:
Locally abundant in Cork Harbour in the 1960s (D. Minchin, pers. ob).

Port vicinity #5/14: Derry/Londonderry
Date of the first record: 1958
Population status: Common
References:
Crisp DJ, Southward AJ (1959) The further spread of Elminius modestus in the British Islea to 1959. Journal of the Marine Biological Association of the United Kingdom 64: 625-635.
Briggs RP, (1982) Community structure and growth of Mytilus edulis L. in Lough Foyle. Proceedings of the Royal Irish Academy 82B: 245-259.
Comments:
Initially when found in 1958 only three mature individuals were found. The species has subsequntly become common in the Foyle Estuary by 2012.

Port vicinity #6/14: Dublin
Date of the first record: 1958
Population status: Common
References:
Crisp DJ, Southward AJ (1959) The further spread of Elminius modestus in the British Islea to 1959. Journal of the Marine Biological Association of the United Kingdom 64: 625-635.
O'Riordan RM, Murphy O (2000) Variation in the reproductive cycle of Elminius modestus in southern Ireland. Journal of the Marine Biological Association of the United Kingdom 80: 608-616.
Comments:
The species is widely distributed in the vicinity of Dublin Port in Dublin Bay, Howth and Dun Laoghaire.

Port vicinity #7/14: Dun Laoghaire
Date of the first record: 1995
Population status: Established
References:
O'Riordan RM (1996) The current status and distribution of the Australian barnacle Elminius modestus Darwin in Ireland. In: Proceedings of the Irish Marine Science Symposium 1995 (eds) Keegan BF and R O'Connor, Galway University Press, Galway, 207-218.
Comments:
Two specimens were found in 1995 but the species has become more frequent since then.

Port vicinity #8/14: Dundalk
Date of the first record: 1958
Population status: Established
References:
Crisp DJ, Southward AJ (1959) The further spread of Elminius modestus in the British Islea to 1959. Journal of the Marine Biological Association of the United Kingdom 64: 625-635.
Comments:
When first found few individuals were found.

Port vicinity #9/14: Fenit
Date of the first record: 1975
Population status: Established
References:
McGrath D and King P (1992) The immigrant barnacle, Elminius modestus Darwin, in Galway Bay, Ireland. Bulletin of the Irish Biogeographical Society 15: 63-67.
Comments:
The first siting was reported from near Blennerville Bridge in the mid 1970s by Colm B Duggan.

Port vicinity #10/14: Foynes
Date of the first record: 1989
Population status: Established
References:
McGrath D and King P (1992) The immigrant barnacle, Elminius modestus Darwin, in Galway Bay, Ireland. Bulletin of the Irish Biogeographical Society 15: 63-67.
Comments:
Specimens were found at Tarbert downstream from Foynes in the Shannon Estuary.

Port vicinity #11/14: Galway
Date of the first record: 1989
Population status: Common
References:
McGrath D, King PA (1992) The immigrant barnacle, Elminius modestus Darwin, in Galway Bay, Ireland. Bulletin of the Irish Biogeographic Society 15: 63-67.
King PA, Keogh E, McGrath D (1997) The current status of the exotic barnacle Elminius modestus Darwin in Galway Bay, Ireland. Irish Naturalists' Journal 25(10): 365-369.
Comments:
By 1996 the species will have become locally common in Galway Bay close to the port region.

Port vicinity #12/14: Killybegs
Date of the first record: 1995
Population status: Established
References:
O'Riordan RM (1996) The current status and distribution of the Australian barnacle Elminius modestus Darwin in Ireland. In: Proceedings of the Irish Marine Science Symposium 1995 (eds) Keegan BF and R O'Connor, Galway University Press, Galway, 207-218.
Comments:
O'Riordan estimated that 15% of the population were breeding. The population being well established by that time.

Port vicinity #13/14: Larne
Date of the first record: 1958
Population status: Established
References:
Crisp DJ, Southward AJ (1959) The further spread of Elminius modestus in the British Islea to 1959. Journal of the Marine Biological Association of the United Kingdom 64: 625-635.

Port vicinity #14/14: Wicklow
Date of the first record: 1995
Population status: Established
References:
O'Riordan RM (1996) The current status and distribution of
the Australian barnacle Elminius modestus Darwin in Ireland. In: Proceedings of the Irish Marine Science Symposium 1995 (eds) Keegan BF and R O'Connor, Galway University Press, Galway, 207-218.
Comments:
Rare in 1995.
 
Created byDan Minchin 
Last update byDan Minchin, 2013-12-06
Contributors
Added by Dan Minchin
Edited by Dan Minchin, 2013-10-11
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Edited by Dan Minchin, 2013-11-04
Edited by Dan Minchin, 2013-12-06